When I first got to my Army NG unit, my Platoon Sgt was ex-Navy. There were quite a few former Marines- including my Company First Sgt. A member of my squad was ex-Coast Guard, and The First Sgt. of another Company was an ex-Seal. It was an Infantry Unit (A rare thing in the reserves these days.) and presumably, everyone in every branch of service had "basic soldiering" as part of their IET. There was quite a bit of MOS specific training conducted for everyone, so getting troops from other branches up to speed was a matter of course. Lots of OJT. (Wow, I've been out for almost ten years and I can still speak in acronyms. For those not fluent in alphabet soup, follows a glossary of terms.)
NG-National Guard
Sgt.- Seargent
IET- Initial Entry Training (aka "Basic")
MOS- Military Occupational Specialty (Job)
OJT- On-the Job-Training (Learn as you go)
Sometimes, a recruit from a different active duty branch would spend their first "summer training" session going to an MOS qualifier school instead of training with the rest of the unit. A former Navy Mechanic joining an Army Reserve Medical unit, for example, would most likely be sent to AIT (advanced individual training) to learn to be a medic. This didn't happen much in my unit, because, I believe the reasoning was, that anyone could be a grunt.