It's in teh manual, but I don't feel like diggin mine out. IIRC, you open it only after getting to a certain depth (quite deep). It's function is to allow the helium that penetrates the watch under pressure at great depth to escape while you're ascending. As you rise and the external pressure eases, if the helium isn't allowed to escape, the internal pressure built up in the watch will pop out the crystal. If you're diving that deep, it's going to take you long enough to decompress coming up that you'll have plenty of time to open the valve. I don't know of any watches rated for this depth that have an auto helium release valve.